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In the 2030s Earth has been overrun by the Blue, which are mutated insect-like creatures that kill and harvest humans for food. Most of the surviving human race has moved to Second Earth, a huge spaceship that orbits the planet. Yuji and Marlene must work together to reach Second Earth and fight against the Blue.
While fighting an intense inter-galactic war, a mecha pilot is accidentally warped into a space-time neither he nor his computer recognize. After waking up from a long-time hibernation, he finds himself trapped on a planet, with human residents talking in an unknown form of language, using inferior technologies, and - most shocking to him - naturally breathable air.
Many years have passed since humanity was driven to the brink of extinction by the sudden emergence of the unknown life forms Gadoll. Those humans that survived now dwell in a 3000m-high mobile fortress Deca-dence built to protect themselves from the Gadoll threat. Denizens of Deca-dence fall into two categories: Gears, warriors who fight the Gadoll daily, and Tankers, those without the skills to fight. One day, Natsume, a Tanker girl who dreams of becoming a Gear meets surly Kaburagi, an armor repairman of Deca-dence. This chance meeting between the seemingly two opposites, the girl with a positive attitude who never gives up on her dreams and the realist who has given up on his, will eventually shake the future course of this world.
In the year 2024, the world has collapsed. Grotesque monsters lurk amongst the ruins of Japan, while remaining people scrape together what they can to survive. Kiruko, an odd-job girl in Nakano, accepts a mysterious woman's dying wish to take a boy named Maru to a place called Heaven. Maru is convinced that there will be a boy there who looks exactly like him.
In the near future, humanity has taken its first steps towards journeying into the far reaches of the galaxy. Upon doing so they discover a huge race of insectoid aliens known as "Space Monsters." These aliens seem dedicated to the eradication of mankind as they near closer and closer to discovering Earth. In response, humanity develops giant fighting robots piloted by hand-picked youth from around the world. Shortly after the discovery of the aliens, Noriko Takaya, the daughter of a famous deceased space captain, enters a training school despite her questionable talents as a pilot. There, she meets her polar opposite, the beautiful and talented Kazumi Amano, and is unexpectedly made to work together with her as they attempt to overcome the trauma of war as well as their own emotions.