Brings Shingo face to face against Yagyu Tajima, the Shogun’s fencing instructor in a match that could save a domain near Osaka. Meanwhile Shogun Yoshimune must face a painful decision whether or not he can finally see his son for the first time.
A showdown decades in the making brings the Donovan family legacy full circle. As the events that made Ray who he is today finally come to light, the Donovans find themselves drawn back to Boston to face the past. Each of them struggles to overcome their violent upbringing, but destiny dies hard, and only their fierce love for each other keeps them in the fight.
After years on the road establishing his reputation as Japan's greatest fencer, Takezo returns to Kyoto. Otsu waits for him, yet he has come not for her but to challenge the leader of the region's finest school of fencing. To prove his valor and skill, he walks deliberately into ambushes set up by the school's followers.
Following the death of the second Tokugawa shogun, it is revealed that he was poisoned by retainers of his son Iemitsu in hopes of gaining him the shogunate despite the stammer and birthmark which undermine his respect. Iemitsu and his brother Tadanaga become bitter rivals for the shogunate, and the land is split into factions, eventually erupting into warfare. Iemitsu's mentor, his fencing instructor Yagyu, is fixated upon securing Iemitsu the shogunate and ends up betraying everyone, even his own family, in pursuit of the goal.
In 1879, Kenshin and his allies face their strongest enemy yet: his former brother-in-law Enishi Yukishiro and his minions, who've vowed their revenge.